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Over time, cacti literally reshaped themselves to fit their environment, by increasing their bulk and reducing the surface area they expose to the sun. Cacti stopped bearing leaves; they could no longer afford to supply broad-surfaced appendages with water. When the cacti dispensed with foliage, their stems and branches took over the leaves’ work of manufacturing food.

The word cacti is __ line 2. Which preposition should I use to fill in, in or on?
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in
Comments  
Over time, by increasing their bulk and reducing the surface area they expose to the sun, Cacti literally reshaped themselves to fit their environment: Cacti stopped bearing leaves; they could no longer afford to supply broad-surfaced appendages with water. When the cacti dispensed with foliage, their stems and branches took over the leaves’ work of manufacturing food.

Active voice is good.

On page 24, you will find the word Cacti in the second sentence.
The line is on a page.
The words are in the line.
 CalifJim's reply was promoted to an answer.